The Roles Of Vitamin D Binding Protein In Human Immune Function

 The scientific and clinical experience with VDBP and
Abstract:Gc-MAF are very encouraging.  Several clinical trials
The recent proliferation of published studies outliningare underway in the Bahamas at the Immune
the role of vitamin D in the prevention of manyAugmentation Therapy Centre to reproduce and
diseases associated with a weakened immuneconfirm the current clinic studies, as well as to
system has brought to light the importance ofanswer several clinical questions that have not yet
monitoring the serum levels of 25(OH) vitamin D. (1) been reported in the scientific literature about VDBP
Specifically, the direct correlation of vitamin D levels inand Gc-MAF.  If you would like to enrol any clients
the human serum with increased levels of cathelicidinwith cancer or immune suppression in a trial with
and the potentiating role cathelicidin plays in theGc-MAF, please contact the author of this review. 
immune response to infections, cancer, autoimmuneCitations:
disease, and especially acute viral infections. (2) 1. Bikie DD, "Vitamin D and immune function:
While research into vitamin D needs to continue, theunderstanding common pathways" Curr Osteoporos
importance of vitamin D binding protein (VDBP) hasRep. 2009 Jul;7(2):58-63
been demonstrated to have synergistic yet2. Yuk JM, Shin DM, Lee HM, Yang CS, Jin HS, Kim
independent functions in the human immuneKK, Lee SH, Kim JM, Jo EK, "Vitamin D3 induces
system.(3)  This article will outline the emerging roleautophagy in human monocytes/macrophages via
of VDBP in the field of immunology.cathelicidin" Cell Host Microbe. 2009 Sep 17;6(3):231-43
Method:3. Chkri M et al, "Production of human macrophages
A review of the scientific literature pertaining towith potent antitumoral properties (MAK) by culture
vitamin D binding protein and its derivatives identifiedof monocytes in the presence of GM-CSF and
in human serum and produced in the laboratory. 1,25-dihydroxy vitamin D3." Anticancer Res. 1992
Results:Nov-Dec;12(6B):2257-60
Vitamin D binding protein, also known as Gc-protein, is4. Christiansen M et al, "Protein chemical
a group of isoform proteins with O-linked glycans. characterization of Gc-gobulin (vitamin D-binding
The dominant isoform of VDBP are non-glycosylatedprotein) isoforms; Gc-f1, Gc-s, and Gc-2." Biochem
656 Da proteins produced mainly in the liver.(4) Biophys Acta. 2007 Apr; 1774(4):481-92
Vitamin D binding protein participates in liver cell5. Gressner OA et al, "Gc-globulin (vitamin D binding
stability and regeneration through Calcium dependentprotein) is synthesized and secreted by hepatocytes
interaction with the megalin/gp330 receptor.(5) and internalized by hepatic stellate cells through Ca2+
There are 4 important roles VDBP has in humans.  Itdependent interaction with megalin/gp330 receptor."
binds circulating vitamin D for transport and storage,Clinica ChimiceActa 2008 Apr 390;1-2:28-37
it is the most important scavenger of extracellular6. Nagasawa H et al, "Gc-protein (vitamin D-binding
G-actin, it enhances the chemotactic activity of C5aprotein): Gc genotyping and GcMAF precursor
for neutrophils in inflammation, and it activatesactivity." Anticancer Res. 2005 Nov-Dec;
macrophages thru GaINAc-modified Gc-protein. (6) 25(6A):3689-95
Additionally, low levels of VDBP have been found to7. Jorgensen CS et al, "Large-scale purification and
correlate with multiple organ failure sepsis, andcharacterization of non-glycosylated Gc-protein
non-survival in fulminant liver failure and traumatic liver(vitamin D binding protein) from plasma fraction IV."
failure.(7)  The non-glycosylated isoform of VDBP isBiotechnol Appl Biochem. 2006 Apr;44(pt1):35-44
able to mask the presence of endotoxins by8. Zella LA et al, "Vitamin D-binding protein influences
20%.(7)  Therapy with VDBP may increase survivaltotal circulating levels of 1,25-dihydroxyvitamin D3 but
in trauma, sepsis and fulminant liver failure.(7) does not directly modulate the bioactive levels of the
The function of VDBP is independent of the hormonehormone in vivo." Endocrinology. 2008
actions of 1,25(2OH) vitamin D and it has limitedJul;149(7):3656-67
impact on the extracellular pool of 1,25(OH) vitamin9. Winters SJ et al, "Influence of obesity on vitamin
D.(8)  Vitamin D binding protein has virtually noD-binding protein and 25-hydroxy vitamin D levels in
impact on the distribution, uptake, activation profile,African and white women." Metabolism. 2009
or biological potency of the hormone vitamin D.(8) Apr;58(4):438-42
Vitamin D binding protein is not affected by race or10. Schiodt FV et al, "Increased turnover of
adiposity the way vitamin D levels are affected.(9) Gc-protein in patients with hepatic encephalopathy."
The serum levels of VDBP are decreased by trauma,Scand J Gastroenterol. 2001 Sep;36(9):998-1003
septic infections, and chronic or acute liver11. Schiodt FV et al, "Gc-globulin and prognosis in
diseases.(10)  The normal levels of serum VDBP areacute liver failure." Liver Transpl. 2005
350-50 mg/L and low levels below 80 mg/L yield aOct;11(10):1223-7
positive and negative mortality predictive value 85%12. Homma S et al, "Vitamin D-binding protein
and 43% respectively.(11) (group-specific component) is the sole serum protein
While vitamin D binding protein is a primaryrequired for macrophage activation after treatment
macrophage activating factor, several glycosylatedof peritoneal cells with lyso-phosphatidylcholine."
isoforms have more potent and specific macrophageImmunol Cel Biol. 1993 Aug;71(pt4):249-57
activating properties.(12)  The most potent serum13. Yamamoto N et al, "Conversion of vitamin D3
macrophage activation factor (MAF) is produced by abinding protein (group-specific component) to a
series of glycosylation reactions performed by themacrophage activating factor by stepwise action of
B-cells and T-cells.(13)  Vitamin D-MAF as a potentbeta-galactosidase of B-celss and sialidase of T-cells."
adjuvant activity for immunization and healthy serumJ Immunol. 1993 Sep 1;151(5):2794-802
levels of MAF prevent tumours from being able to14. Yamamoto N et al, "Structurally well-defined
transplant into mice.(14)  Other roles for VDBPmacrophage activating factor derived from vitamin
derived MAF have been described, including anD3-binding protein has a potent adjuvant activity for
anti-angiogenesis function through blockingimmunization." Immunol Cell Biol. 1998 Ju;76(3):237-44
VEGF-induced angiogenesis.(15) 15. Kallunte S et al, "Inhibition of angiogenesis by
Laboratory derived MAF from serum VDBP has thevitamin D-binding protein: characterization of
advantage of activating macrophages and not beinganti-endothelial activity of DBP-MAF."  Angiogenesis.
deglycosylated by N-acetylgalactosaminidase2005;8(4):349-60
enzymes (NaGaLase) produced by cancer cells and16. Yamamoto N et al, "Immunotherapy of
infectious bacteria, viruses and fungi.(16)  The clinicalmetastatic colorectal cancer with vitamin D-binding
dose required to have a sustained systemic activationprotein derived macrophage activating factor,
of macrophages in 100 mcg injected weekly.(16)  AGc-MAF." Cancer Immunol Immunotherapy
novel MAF (Gc-MAF) can be produced in the lab with2008;57:1007-16
cancer specific activity that targets undifferentiated17. Yamamoto N et al, "Immunotherapy of
cancer cells better than well differentiated cancermetastatic breast cancer patients with vitamin
cells.(17)  Several prospective clinical trials ofD-bindng protein-derived macrophage activating
treatment of cancer and HIV with Gc-MAF havefactor (GcMAF)." Int J Cancer. 2008;122:461-467
been reported and three trials with metastatic colon,18. Yamamoto N, Suyama H, Yamamoto N
breast and prostate cancer have provided 100%"Immunotherapy for prostate cancer with Gc
remission rate beyond the five years since Gc-MAFprotein-derived macrophage activating factor,
treatment.(16)(17)(18)GcMAF." Transl Oncol.
Discussion: